草业学报 ›› 2022, Vol. 31 ›› Issue (3): 181-191.DOI: 10.11686/cyxb2020589
• 研究论文 • 上一篇
刘启宇1(), 云岚1,2(), 陈逸凡1, 郭宏宇1, 李珍1, 高志琦1, 王俊1, 石凤翎1,2
收稿日期:
2020-12-29
修回日期:
2021-02-26
出版日期:
2022-03-20
发布日期:
2022-01-15
通讯作者:
云岚
作者简介:
Corresponding author. E-mail: yunlan@imau.edu.cn基金资助:
Qi-yu LIU1(), Lan YUN1,2(), Yi-fan CHEN1, Hong-yu GUO1, Zhen LI1, Zhi-qi GAO1, Jun WANG1, Feng-ling SHI1,2
Received:
2020-12-29
Revised:
2021-02-26
Online:
2022-03-20
Published:
2022-01-15
Contact:
Lan YUN
摘要:
为了解在内蒙古中部轻度盐碱地上建植豆禾混播草地的牧草产量及各混播组合物种的竞争关系,以草原3号苜蓿分别与新麦草、缘毛雀麦和长穗偃麦草3种禾草以不同比例(1∶1,1∶2,1∶3)混播建植人工草地,研究不同刈割期、不同混播方式下的牧草产量、土地当量比(LER)、相对产量(RY)、相对密度(RD)、相对产量总值(RYT)和种间竞争率(CR)等。结果表明,混播物种、混播比例和刈割时期对混播草地的产量和种间竞争关系有直接影响。在3个刈割时期,所有混播组合的牧草产量均高于单播,且土地当量比均大于1。相对产量、相对密度和相对产量总值除个别1∶3组合外也均大于1,根据物种竞争模型对混播组合中各物种竞争关系进行分析,发现除豆禾1∶3组合外,苜蓿与3种禾草均分布在双方受益区域,表明各草种混播组合基本呈共存状态。除苜蓿与缘毛雀麦1∶1组合外,苜蓿在其余各混播组合中种间竞争率均大于 1,且在同一刈割期随禾草比例的升高而增大。从牧草产量和种间关系的稳定性综合考虑,在上述地区苜蓿与缘毛雀麦以1∶1或1∶3混播,苜蓿与新麦草、苜蓿与长穗偃麦草以1∶1或1∶2混播更有利于建立高产稳定的人工草地。
刘启宇, 云岚, 陈逸凡, 郭宏宇, 李珍, 高志琦, 王俊, 石凤翎. 苜蓿—禾草混播草地牧草产量及种间竞争关系的动态研究[J]. 草业学报, 2022, 31(3): 181-191.
Qi-yu LIU, Lan YUN, Yi-fan CHEN, Hong-yu GUO, Zhen LI, Zhi-qi GAO, Jun WANG, Feng-ling SHI. The dynamic analysis of forage yield and interspecific competition in alfalfa-grass mixed pasture[J]. Acta Prataculturae Sinica, 2022, 31(3): 181-191.
编号 Code | 混播处理 Mixed treatments | 行数 Row number | 混播比例 Mixed sowing ratio | 每行播种量 Sowing rate of each line (g) |
---|---|---|---|---|
A1B1 | 草原3号苜蓿M. varia-新麦草P. juncea | 4~4 | 1∶1 | 6.444~8.192 |
A1B2 | 草原3号苜蓿M. varia-新麦草P. juncea | 3~6 | 1∶2 | 4.833~12.288 |
A1B3 | 草原3号苜蓿M. varia-新麦草P. juncea | 2~6 | 1∶3 | 3.222~12.288 |
A2B1 | 草原3号苜蓿M. varia-缘毛雀麦B. ciliatus | 4~4 | 1∶1 | 6.444~9.976 |
A2B2 | 草原3号苜蓿M. varia-缘毛雀麦B. ciliatus | 3~6 | 1∶2 | 4.833~14.964 |
A2B3 | 草原3号苜蓿M. varia-缘毛雀麦B. ciliatus | 2~6 | 1∶3 | 3.222~14.964 |
A3B1 | 草原3号苜蓿M. varia-长穗偃麦草E. elongata | 4~4 | 1∶1 | 6.444~25.184 |
A3B2 | 草原3号苜蓿M. varia-长穗偃麦草E. elongata | 3~6 | 1∶2 | 4.833~37.776 |
A3B3 | 草原3号苜蓿M. varia-长穗偃麦草E. elongata | 2~6 | 1∶3 | 3.222~37.776 |
C1 | 单播新麦草Monoculture P. juncea | 8 | - | 16.384 |
C2 | 单播缘毛雀麦Monoculture B. ciliatus | 8 | - | 19.952 |
C3 | 单播长穗偃麦草Monoculture E. elongata | 8 | - | 50.368 |
C4 | 单播草原3号苜蓿Monoculture M. varia | 8 | - | 12.888 |
表1 豆禾混播草地混播比例及播量
Table 1 The proportion and sowing rate of mixed alfafa and grass mixed sowing
编号 Code | 混播处理 Mixed treatments | 行数 Row number | 混播比例 Mixed sowing ratio | 每行播种量 Sowing rate of each line (g) |
---|---|---|---|---|
A1B1 | 草原3号苜蓿M. varia-新麦草P. juncea | 4~4 | 1∶1 | 6.444~8.192 |
A1B2 | 草原3号苜蓿M. varia-新麦草P. juncea | 3~6 | 1∶2 | 4.833~12.288 |
A1B3 | 草原3号苜蓿M. varia-新麦草P. juncea | 2~6 | 1∶3 | 3.222~12.288 |
A2B1 | 草原3号苜蓿M. varia-缘毛雀麦B. ciliatus | 4~4 | 1∶1 | 6.444~9.976 |
A2B2 | 草原3号苜蓿M. varia-缘毛雀麦B. ciliatus | 3~6 | 1∶2 | 4.833~14.964 |
A2B3 | 草原3号苜蓿M. varia-缘毛雀麦B. ciliatus | 2~6 | 1∶3 | 3.222~14.964 |
A3B1 | 草原3号苜蓿M. varia-长穗偃麦草E. elongata | 4~4 | 1∶1 | 6.444~25.184 |
A3B2 | 草原3号苜蓿M. varia-长穗偃麦草E. elongata | 3~6 | 1∶2 | 4.833~37.776 |
A3B3 | 草原3号苜蓿M. varia-长穗偃麦草E. elongata | 2~6 | 1∶3 | 3.222~37.776 |
C1 | 单播新麦草Monoculture P. juncea | 8 | - | 16.384 |
C2 | 单播缘毛雀麦Monoculture B. ciliatus | 8 | - | 19.952 |
C3 | 单播长穗偃麦草Monoculture E. elongata | 8 | - | 50.368 |
C4 | 单播草原3号苜蓿Monoculture M. varia | 8 | - | 12.888 |
物种Species | 刈割期Ⅰ Mowing period Ⅰ | 刈割期Ⅱ Mowing period Ⅱ | 刈割期Ⅲ Mowing period Ⅲ |
---|---|---|---|
新麦草P. juncea | 抽穗期Heading stage | 乳熟期Milk ripening stage | 完熟期Full-ripe stage |
缘毛雀麦B. ciliatus | 抽穗期Heading stage | 初花期Early flowering stage | 乳熟期Milk ripening stage |
长穗偃麦草E. elongata | 拔节期Jointing stage | 抽穗期Heading stage | 灌浆期 Filling stage |
草原3号苜蓿M. varia | 初花期Early flowering stage | 盛花期Full-bloom stage | 结荚期Podding stage |
表2 不同刈割期牧草所对应物候期
Table 2 Corresponding phenological periods of forages in different mowing periods
物种Species | 刈割期Ⅰ Mowing period Ⅰ | 刈割期Ⅱ Mowing period Ⅱ | 刈割期Ⅲ Mowing period Ⅲ |
---|---|---|---|
新麦草P. juncea | 抽穗期Heading stage | 乳熟期Milk ripening stage | 完熟期Full-ripe stage |
缘毛雀麦B. ciliatus | 抽穗期Heading stage | 初花期Early flowering stage | 乳熟期Milk ripening stage |
长穗偃麦草E. elongata | 拔节期Jointing stage | 抽穗期Heading stage | 灌浆期 Filling stage |
草原3号苜蓿M. varia | 初花期Early flowering stage | 盛花期Full-bloom stage | 结荚期Podding stage |
图1 两物种竞争试验模式带箭头的斜线表示RYA=RYB,在斜线之上的三角形区域表示物种A具有竞争优势,在斜线之下的三角形区域表示物种B具有竞争优势。表示该区域内物种A和B混播时生长均受到抑制。表示该区域内物种A和B在混播中有明显的促进作用。在RYA=1到x轴的区域内物种B在混播中竞争力强,明显抑制了物种A的生长。在RYB=1到y轴的区域内物种A在混播中竞争力强,明显抑制物种B的生长。The diagonal lines with the arrows in figure means RYA=RYB , the triangle area above the diagonal lines of graph in which species A has a competitive advantage over B, the triangle area below the diagonal lines of graph in which species B has a competitive advantage over A. means the growth of species A and B are suppressed in the area when the mixed seeding,means the growth of species A and B are promoted in the area when the mixed seeding. In the area of RYA=1 to x-axis means species B is competitive over A, in the area of RYB=1 to y-axis means species A is competitive over B, significantly inhibited the growth of B.
Fig.1 Two species competition test model
混播处理 Mixed treatments | 刈割期Ⅰ Mowing period Ⅰ | 刈割期Ⅱ Mowing period Ⅱ | 刈割期Ⅲ Mowing period Ⅲ | ||||||
---|---|---|---|---|---|---|---|---|---|
禾草 Grass | 苜蓿 Alfalfa | 总产量 Total yield | 禾草 Grass | 苜蓿 Alfalfa | 总产量 Total yield | 禾草 Grass | 苜蓿 Alfalfa | 总产量 Total yield | |
A1B1 | 3333.33bc | 6400.00a | 9733.33a | 4333.33a | 12166.67a | 16500.00a | 4666.67a | 6733.33a | 11400.00abc |
A1B2 | 3500.00bc | 5666.67a | 9166.67a | 4733.33a | 13900.00a | 18633.33a | 4266.67a | 9133.33a | 13400.00ab |
A1B3 | 2600.00c | 5433.33a | 8033.33a | 4233.33a | 11433.33a | 15666.66a | 5933.33a | 10566.67a | 16500.00a |
A2B1 | 5166.67a | 5000.00a | 10166.67a | 8700.00a | 6300.00a | 15000.00a | 6500.00a | 8800.00a | 15300.00a |
A2B2 | 4766.67ab | 3700.00a | 8466.67a | 5566.67a | 4966.67a | 10533.34ab | 6533.33a | 6600.00a | 13133.33ab |
A2B3 | 4633.33ab | 4700.00a | 9333.33a | 7200.00a | 7933.33a | 15133.33a | 6433.33a | 8000.00a | 14433.33ab |
A3B1 | 2400.00c | 5900.00a | 8300.00a | 5600.00a | 12433.33a | 18033.33a | 8300.00a | 6866.67a | 15166.67a |
A3B2 | 2800.00c | 5500.00a | 8300.00a | 5133.33a | 10033.33a | 15166.66a | 6133.33a | 10966.67a | 17100.00a |
A3B3 | 2900.00c | 4466.67a | 7366.67a | 4000.00a | 7166.67a | 11166.67ab | 7300.00a | 8933.33a | 16233.33a |
C1 | 3266.67bc | - | 3266.67c | 3733.33a | - | 3733.33b | 3866.67a | - | 3866.67d |
C2 | - | - | 4866.67bc | 6066.67a | - | 6066.67b | 6633.33a | - | 6633.33cd |
C3 | 2730.00c | - | 2730.00c | 5500.00a | - | 5500.00b | 5850.00a | - | 5850.00cd |
C4 | - | 4700.00a | 4700.00bc | - | 10266.67a | 10266.67ab | - | 8233.33a | 8233.33bcd |
表3 不同混播处理下不同刈割期的干物质产量
Table 3 The dry matter yield of different cutting periods under different mixed sowing treatments (kg·hm-2)
混播处理 Mixed treatments | 刈割期Ⅰ Mowing period Ⅰ | 刈割期Ⅱ Mowing period Ⅱ | 刈割期Ⅲ Mowing period Ⅲ | ||||||
---|---|---|---|---|---|---|---|---|---|
禾草 Grass | 苜蓿 Alfalfa | 总产量 Total yield | 禾草 Grass | 苜蓿 Alfalfa | 总产量 Total yield | 禾草 Grass | 苜蓿 Alfalfa | 总产量 Total yield | |
A1B1 | 3333.33bc | 6400.00a | 9733.33a | 4333.33a | 12166.67a | 16500.00a | 4666.67a | 6733.33a | 11400.00abc |
A1B2 | 3500.00bc | 5666.67a | 9166.67a | 4733.33a | 13900.00a | 18633.33a | 4266.67a | 9133.33a | 13400.00ab |
A1B3 | 2600.00c | 5433.33a | 8033.33a | 4233.33a | 11433.33a | 15666.66a | 5933.33a | 10566.67a | 16500.00a |
A2B1 | 5166.67a | 5000.00a | 10166.67a | 8700.00a | 6300.00a | 15000.00a | 6500.00a | 8800.00a | 15300.00a |
A2B2 | 4766.67ab | 3700.00a | 8466.67a | 5566.67a | 4966.67a | 10533.34ab | 6533.33a | 6600.00a | 13133.33ab |
A2B3 | 4633.33ab | 4700.00a | 9333.33a | 7200.00a | 7933.33a | 15133.33a | 6433.33a | 8000.00a | 14433.33ab |
A3B1 | 2400.00c | 5900.00a | 8300.00a | 5600.00a | 12433.33a | 18033.33a | 8300.00a | 6866.67a | 15166.67a |
A3B2 | 2800.00c | 5500.00a | 8300.00a | 5133.33a | 10033.33a | 15166.66a | 6133.33a | 10966.67a | 17100.00a |
A3B3 | 2900.00c | 4466.67a | 7366.67a | 4000.00a | 7166.67a | 11166.67ab | 7300.00a | 8933.33a | 16233.33a |
C1 | 3266.67bc | - | 3266.67c | 3733.33a | - | 3733.33b | 3866.67a | - | 3866.67d |
C2 | - | - | 4866.67bc | 6066.67a | - | 6066.67b | 6633.33a | - | 6633.33cd |
C3 | 2730.00c | - | 2730.00c | 5500.00a | - | 5500.00b | 5850.00a | - | 5850.00cd |
C4 | - | 4700.00a | 4700.00bc | - | 10266.67a | 10266.67ab | - | 8233.33a | 8233.33bcd |
处理 Treatments | 刈割期Mowing period | 处理 Treatments | 刈割期Mowing period | ||||
---|---|---|---|---|---|---|---|
Ⅰ | Ⅱ | Ⅲ | Ⅰ | Ⅱ | Ⅲ | ||
A1B1 | 2.50±0.36a | 2.80±1.18a | 2.15±0.45a | A2B3 | 1.98±0.57a | 2.16±0.82a | 2.07±0.98a |
A1B2 | 2.50±0.71a | 2.97±0.22a | 2.37±0.62a | A3B1 | 2.47±0.98a | 1.97±0.46a | 2.70±0.98a |
A1B3 | 2.08±0.47a | 2.56±0.47a | 3.11±1.04a | A3B2 | 2.12±0.17a | 1.90±0.39a | 2.74±0.38a |
A2B1 | 2.56±0.75a | 2.05±0.20a | 2.15±0.25a | A3B3 | 2.24±0.95a | 1.79±1.01a | 2.58±0.81a |
A2B2 | 1.84±0.20a | 1.52±0.44a | 1.91±0.47a |
表4 混播处理对土地当量比的影响
Table 4 Effect of mixed seeding treatments on land equivalent ratio (LER)
处理 Treatments | 刈割期Mowing period | 处理 Treatments | 刈割期Mowing period | ||||
---|---|---|---|---|---|---|---|
Ⅰ | Ⅱ | Ⅲ | Ⅰ | Ⅱ | Ⅲ | ||
A1B1 | 2.50±0.36a | 2.80±1.18a | 2.15±0.45a | A2B3 | 1.98±0.57a | 2.16±0.82a | 2.07±0.98a |
A1B2 | 2.50±0.71a | 2.97±0.22a | 2.37±0.62a | A3B1 | 2.47±0.98a | 1.97±0.46a | 2.70±0.98a |
A1B3 | 2.08±0.47a | 2.56±0.47a | 3.11±1.04a | A3B2 | 2.12±0.17a | 1.90±0.39a | 2.74±0.38a |
A2B1 | 2.56±0.75a | 2.05±0.20a | 2.15±0.25a | A3B3 | 2.24±0.95a | 1.79±1.01a | 2.58±0.81a |
A2B2 | 1.84±0.20a | 1.52±0.44a | 1.91±0.47a |
图2 不同混播处理下不同刈割期牧草相对产量和相对密度的比较
Fig.2 Comparison of forage relative yield and relative density in different cutting periods under different mixed sowing treatments
图3 不同混播处理下不同刈割期禾草与苜蓿竞争结果分布
Fig.3 Distribution of competition results between grass and alfalfa in different cutting periods under different mixed sowing treatments
图5 不同混播处理下不同刈割期禾草与苜蓿的种间竞争率不同小写字母表示各处理间差异显著(P<0.05)。Different lowercase letters indicate significant differences among different treatments (P<0.05).
Fig.5 Interspecific competition rate of grass and alfalfa in different cutting periods under different mixed sowing treatments
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