草业学报 ›› 2014, Vol. 23 ›› Issue (5): 330-337.DOI: 10.11686/cyxb20140539
张文涛1,杜久元1,陈新宏2,武军2,赵继新2,杜万里2,张礼军1
收稿日期:
2014-01-14
出版日期:
2014-10-20
发布日期:
2014-10-20
作者简介:
张文涛(1985-),男,甘肃榆中人,研实员,硕士。E-mail:zhangwt105@126.com
基金资助:
国家引进国外智力成果示范推广(Y20136200001)资助
ZHANG Wen-tao1,DU Jiu-yuan1,CHEN Xin-hong2,WU Jun2,ZHAO Ji-xin2,DU Wan-li2,ZHANG Li-jun1
Received:
2014-01-14
Online:
2014-10-20
Published:
2014-10-20
摘要:
华山新麦草是仅分布于中国秦岭山脉华山段的特有农作物野生近缘物种,具有抗寒、耐旱、耐瘠薄、抗病、优质等优异性状特点。因此,华山新麦草可作为小麦抗病、抗逆等性状遗传改良的重要种质资源,对拓宽小麦遗传基础、创制新种质资源具有重要意义。本文围绕华山新麦草种群生态保护研究及其在小麦遗传改良中的应用现状,阐述了华山新麦草分类及进化、种群生态和小麦遗传改良过程中的应用情况,并针对华山新麦草的生态保护及在小麦育种中的应用前景进行了展望,以期对今后特异种质资源在小麦遗传育种实践中高效利用该种质资源有所启发。
中图分类号:
张文涛,杜久元,陈新宏,武军,赵继新,杜万里,张礼军. 华山新麦草在小麦遗传改良中的应用研究进展[J]. 草业学报, 2014, 23(5): 330-337.
ZHANG Wen-tao,DU Jiu-yuan,CHEN Xin-hong,WU Jun,ZHAO Ji-xin,DU Wan-li,ZHANG Li-jun. Progress report on the study of Psathyrostachys huashanica and its application to wheat improvement[J]. Acta Prataculturae Sinica, 2014, 23(5): 330-337.
Reference:[1] Li L H, Yang X M, Li X Q, et al. Study and utilization of wild relatives of wheat in China[J]. Review of China Agricultural Science and Technology, 2000, 6(2): 73-76. [2] Dewey D R. Historical and current taxonomic perspectives of Agropyron, Elymus and related genera[J]. Crop Science, 1983, 23(4): 637-642.[3] Dewey D R, Hsiao C. A cytogenetic basis for transferring Russian wild rye from Elymus to Psathyrostachys[J]. Crop Science, 1983, 23(1): 123-126.[4] Baden C. A taxonomic revision of Psathyrostachys (Poaceae)[J]. Nordic Journal of Botany, 1991, 11(1): 3-26.[5] Guo B Z. Flora of China (ninth volume Volume III)[M]. Beijing: Science Press, 1987.[6] Dewey D R. The genome system of classification as a guide to intergeneric hybridization with the perennial Triticeae[A].In: Gustafson J P. Gene Manipulation in Plant Improvement[M]. New York: Plenum Press, 1984: 209-279.[7] Hsiao C, Richard R, Wang C, et al. Karyotype analysis and genome relationships of 22 diploid species in the tribe Triticeae[J]. Canadian Journal of Genetics and Cytology, 1986, 28(1): 109-120.[8] Linde-Laursen I, von Bothmer R. Comparison of the karyotype of Psathyrostachys juncea and P. huashanica (Poaceae) studied by banding techniques[J]. Plant Systematics and Evolution, 1986, 151(3): 203-213.[9] Wang R C. Synthetic and natural hybrids of Psathyrostachys huashanica[J]. Genome, 1987, 29(6): 811-816.[10] Wang X E, Li W L, Liu D D. C-banding of two Psathyrostachys species[J]. Journal of Nanjing Agricultural University, 1998, 21(1): 10-13.[11] Sun G L, Yan J, Yang J L. A biosystematic study on hybrids of psathyrostachys huashanica with Roegneria caucasica and Kengyilia hirsuya[J]. Acta Pratacultural Science, 1995, 4(4): 39-45.-[12] Liu F, Sun G L, Yana J, et al. Study of electrophoresis of isozymes of Triticum aestivum Psathyrostachys huashanica and their Hybrid F1[J]. Acta Agronomica Sinica, 1992, (3): 169-175.[13] Zhang H Q, Zhou Y H. Meiotic pairing behaviour reveals differences in genomic constitution between Hystrix patula and other species of the genus Hystrix Moench (Poaceae, Triticeae)[J]. Plant Systematics and Evolution, 2006, 258(3): 129-136.[14] Zhang H Q, Yang R W, Dou Q W, et al. Genome constitutions of Hystrix patula, H. duthiei ssp. duthiei and H.duthiei ssp. longearistata (Poaceae: Triticeae)revealed by meiotic pairing behavior and genomic in-situ hybridization[J]. Chromosome Research, 2006, 14(6): 595-604.[15] Wang L, Yang J, Guo J, et al. Genetic structure and differentiation of Psathyrostachys huashanica populations detected with RAPD markers[J]. Acta Ecologica Sinica, 2005, 25(4): 720-727.[16] Zhao L F, Li S, Pan Y, et al. Population differentiation of Psathyrostachys huashanina along an altitudinal gradient detected by random amplified polymorphic DNA[J]. Acta Botanica Boreali-Occidentalia Sinica, 2001, 21(3): 391-400.[17] Liu Z L, Li S, Yan G Q, et al. Genetic Structure and Intraspecific Genetic Polymorphisms in Natural Populations of Psathyrostachys huashanica[J]. Acta Genetica Sinica, 2001, 28(8): 769-777.[18] Liu Z L, Song Y, Li S, et al. Pollination phenology and an indirect estimate of gene flow in natural populations of psathyrostachys huashanica[J]. Acta Phytoecologica Sinica, 2001, 25(4): 426-430.[19] Yue M, Zhang L J, Ma K, et al. Endangering reason and reproductive strategy of Psathyrostachys huashanica population[J]. Acta Ecologica Sinica, 2001, 21(8): 1314-1320.[20] Wang L, Zhao G F. Studies on the development of embryo and endosperm in Psathyrostachys huashanica[J]. Acta Botanica Boreali-Occidentalia Sinica, 2002, 22(6): 786-790.[21] Zhang L J, Wang L, Li Z X, et al. The microsporogenesis and male gamephyte development of psathyrostachys huashanica Keng[J]. Journal of Northwest University(Natural Science Edition), 2002, 32(2): 77-80.[22] Zeng L, Yuan Q H, Wang F, et al. Genetic diversity analysis of Agropyron germplasm resources by ISSR[J]. Acta Prataculturae Sinica, 2013, 22(1): 260-267.[23] Li J Q, Wang L H, Yan Q W, et al. Genetic diversity of 20 ryegrass accessions by SRAP markers[J]. Acta Prataculturae Sinica, 2013, 22(2): 158-164.[24] Huang X H, Hu X W, Xu Z H, et al. Effects of exogenous hormones on the dormancy and germination of Leymus chinensis[J]. Acta Prataculturae Sinica, 2013, 22(5): 183-189.[25] Wang M N, Shang H S. Evaluation of resistance in Psathyrostachys huashaica to wheat take-all fungus[J]. Acta Universitatis Agriculturalis Boreali-Occidentalis, 2000, 28(6): 69-71.[26] Jing J X, Fu X, Yuan H X. A preliminary study on heredity of the resistance to stripe rustin three wild relatives of wheat[J]. Acta Phytopathologica Sinica, 1999, 2(2): 147-150.[27] Wan Y F, Yan J, Yang J L, et al. Study on wild relatives of wheat for resistance to head SCAB[J]. Acta Phytopathologica Sinica, 1997, 27(2): 107-111.[28] Li Q, Ye H Z. Studies on Resistance of Wild Relatives in Triticeae to Oat Bird-cherry Aphids (Homoptera: Aphididae)[J]. Scientia Agricultura Sinica, 2002, 35(6): 719-723.[29] Song S, Tao Y, Zhang H W, et al. Psathyrostachys huashanica, a potential resource for resistance to Barley yellow dwarf virus-GAV[J]. European Journal of Plant Pathology, 2013, 137: 217-221.[30] Petersen G, Seberg O, Baden C. A phylogenetic analysis of the genus Psathyrostachys (Poaceae) based on one nuclear gene, three plastid genes, and morphology[J]. Plant Systematics and Evolution, 2004, 249: 99-110.[31] Fan X, Zhang Y, Zhou Y H. Sequence Analysis of Gene ccmFN Encoding N - terminal Maturation Subunit F of Cytochromes c in Psathyrostachys huashanica and Pseudoroegneria strigosa ( Poaceae : Triticeae)[J]. Journal of Sichuan Agricultural University, 2006, 24(3): 252-255.[32] Dong Y C. Wheat distant hybridization breeding[A]. Wheat genetics and breeding prospects in the 21st century - Wheat Genetics and Breeding International Symposium Proceedings[C]. Beijing: China Agricultural Science and Technology Press, 2001: 12-16.[33] Chen S Y, Zhang A J, Fu J. The hybridization between Triticum aestivum and Psathyrostachys huashanica[J]. Acta Genetica Sinica, 1991, 18(6): 508-512.[34] Chen S Y, Hou W S, Zhang A J, et al. Breeding and cytogenetic study of Triticum aestivum-Psath yrostachys huashanica alien addition lines[J]. Acta Genetica Sinica, 1996, 23(6): 447-452.[35] Sun G L, Yan J, Yang J L. Production and cytogenetic study of intergeneric hybrid between Triticum aestivum and Psathyostachys specie[J]. Acta Genetica Sinica, 1992, 19(4): 322-326.[36] Hou W J, Zhang A J, Yang H Q, et al. Breeding and cytogenetic study of Triticum aestivum-Psathyrostachys huashanica alien substitution lines[J]. Acta Botanica Boreali-Occidentalia Sinica, 1997, 17(3): 368-373.[37] Kishii M, Dou Q, Garg M, et al. Production of wheat Psathyrostachys huashanica chromosome addition lines[J]. Genes & Genetic Systems, 2010, 85(4): 281-286.[38] Wang Y, Xie Q, Yu K F, et al. Development and characterization of wheat Psathyrostachys huashanica partial amphiploids for resistance to stripe rust[J]. Biotechnology Letters, 2011, 33(6): 1233-1238.[39] Wang Y, Kang H Y, Zhang H Q, et al. Distribution of chromosome numbers in BC2 and BC 1F1 progenies derived from Triticum aestivum×Psathyrostachys huashanica[J]. Caryologia, 2011, 64(1): 67-74.[40] Wang Y, Yu K F, Xie Q, et al. Cytogenetic, genomic in situ hybridization (GISH) and agronomic characterization of alien addition lines derived from wheat Psathyrostachys huashanica[J]. African Journal of Biotechnology, 2011, 10(12): 2201-2211.[41] Kang H Y, Zhang H Q, Fan X, et al. Morphological and cytogenetic studies on the hybrid between bread wheat and Psathyrostachys huashanica Keng ex Kuo[J]. Euphytica, 2008, 162(3): 441-448.[42] Kang H Y, Zhang H Q, Wang Y, et al. Comparative analysis of the homoeologous pairing effects of phKL gene in common wheat×Psathyrostachys huashanica Keng ex Kuo[J]. Cereal Research Communications, 2008, 36(3): 429-440.[43] Kang H Y, Wang Y, Sun G L, et al. Production and characterization of an amphiploid between common wheat and Psathyrostachys huashanica Keng ex Kuo[J]. Plant Breeding, 2009, 128(1): 36-40.[44] Kang H Y, Chen Q, Wang Y, et al. Molecular cytogenetic characterization of the amphiploid between bread wheat and Psathyrostachys huashanica[J]. Genetic Resources and Crop Evolution, 2010, 57(1): 111-118.[45] Kang H Y, Zhong M Y, Xie Q, et al. Production and cytogenetics of trigeneric hybrid involving Triticum, Psathyrostachys and Secale[J]. Genetic Resources and Crop Evolution, 2012, 59(3): 445-453.[46] Kang H Y, Zeng J, Xie Q, et al. Molecular cytogenetic characterization and stripe rust response of a trigeneric hybrid involving Triticum, Psathyrostachys, and Thinopyrum[J]. Genome, 2012, 55(5): 383-390.[47] Zhao J X, Chen X H, Wang X L, et al. Molecular cytogenetic study on the alien substitution lines of Triticum-Psathyrostachys[J]. Acta Botanica Boreali-Occidentalia, 2004, 24(12): 2277-2281.[48] Wu J, Zhao J X, Chen X H, et al. Cytology Characteristic and GISH Analysis on the Progenies Derived from Common Wheat (T. aestivun L.)×Psathyrostachys huashanica[J]. Journal of Triticeae Crops, 2007, 27(5): 772-775.[49] Wu J, Zhao J X, Chen X H, et al. SSR analysis on wheat (Triticum aestivum L.) -Psathyrostachys huashanica alien addition lines[J]. Journal of Agricultural University of Hebei, 2007, 30(5): 9-13.[50] Du W L, Wang J, Wang L M, et al. A novel SCAR marker for detecting Psathyrostachys huashanica Keng chromatin introduced in wheat[J]. Genetics and Molecular Research, 2013, 12(4): 4797-4806.[51] Zhao J X, Wu J, Cheng X N, et al. Agronomic and quality traits of a wheat-psathyrostachys huashanica 1Ns disomic addition line[J]. Acta Agronomica Sinica, 2010, 36(9): 1610-1614.[52] Kang H Y, Wang Y, Fedak G, et al. Introgression of chromosome 3Ns from Psathyrostachys huashanica into wheat specifying resistance to stripe rust[J]. PLoS ONE, 2011, 6(7): e21802. [53] Wang Y, Yu K, Xie Q, et al. The 3Ns chromosome of Psathyrostachys huashanica carries the gene(s) underlying wheat stripe rust resistance[J]. Cytogenetic and Genome Research, 2011, 134(2): 136-143.[54] Du W L, Wang J, Lu M, et al. Characterization of a wheat Psathyrostachys huashanicaKeng 4Ns disomic addition line for enhanced tiller numbers and stripe rust resistance[J]. Planta, 2013, 239(1): 97-105. [55] Du W L, Wang J, Lu M, et al. Molecular cytogenetic identification of a wheat Psathyrostachys huashanica Keng 5Ns disomic addition line with stripe rust resistance[J]. Molecular Breeding, 2013, 31(4): 879-888.[56] Du W L, Wang J, Pang Y H, et al. Isolation and characterization of a Psathyrostachys huashanica Keng 6Ns chromosome addition in common wheat[J]. PLoS One, 2013, 8: e53921.[57] Du W L, Wang J, Wang L M, et al. Development and characterization of a Psathyrostachys huashanica Keng 7Ns chromosome addition line with leaf rust resistance[J]. PLoS One, 2013, 8: e70879.[58] Li Z Q, Zeng S M. Chinese wheat rust[M].Beijing: China Agriculture Press, 2002: 370-373.[59] Wang D B, Wang M N, Jing J X, et al. Study of resistance in distant hybridigation descendant of wheat to the take-all[J]. Acta Botanica Boreali-Occidentalia Sinica, 2003, 23(9): 1617-1620.[60] Fu J, Wang M N, Zhao J X, et al. Studies on cytogenetics and utilization of wheat-Psathyrostachys huashanica medium material H8911 with resistance to wheat take-all fungus[J]. Acta Botanica Boreali-Occidentalia Sinica, 2003, 23(12): 2157-2162.[61] Fu J, Zhao J X, Chen S Y, et al. Studies on molecular cytogenetics of wheat-Psathyrostachys huashanica germplasm with resistance to wheat take-all fungus[J]. Acta Botanica Boreali-Occidentalia Sinica, 2003, 23(11): 1905-1909.[62] Wei F Q, Wu J, Zhao J X, et al. Genetic analysis of resistance to take-all fungus of wheat line H9021 derived from wheat-psathyrostachys huashanica[J]. Journal of Triticeae Crops, 2009, 29(1): 153-156.[63] Hang Y, Jin Y, Lu B R. Genetic diversity of the endangered species psathyrostachys huashanicain China and its strategic conservation[J]. Journal of Fudan University(Natural Science), 2004, 43(2): 260-266.[64] Liu P, Yang M N, Zhou X L, et al. Genetic analysis and molecular mapping of stripe rust resistance of wheat translocation line H9020-1-6-8-3 derived from Psathyrostachys huashanica Keng[J]. Acta Phytopathologica Sinica, 2008, 38(1): 104-107.[65] Cao Z J, Wang X P, Wang M N, et al. Genetic analysis and molecular markers of a novel stripe rust resistance gene YrHua in wheat originated from psathyrostachys huashanica Keng[J]. Acta Genetica Sinica, 2005, 32(7): 738-743.[66] Ding C B, Zhou Y H. Spatial pattern and sampling technique of the root rot on Chinese prickly ASH[J]. Journal of Sichuan Agricultural University, 1997, 15(1): 18-20.[67] Liao J Q, Kang H Y, Yang R W, et al. Cytogenetics study of Chinese Spring ph2b mutant ×Psathyrostachys huashanica Keng Selfing(S1) and Backcross(BC1)[J]. Acta Botanica Boreali-Occidentalia Sinica, 2007, 27(3): 442-448. 参考文献:[1] 李立会, 杨欣明, 李秀全, 等. 中国小麦野生近缘植物的研究与利用[J]. 中国农业科技导报, 2000, 6(2): 73-76. [2] Dewey D R. Historical and current taxonomic perspectives of Agropyron, Elymus and related genera[J]. Crop Science, 1983, 23(4): 637-642.[3] Dewey D R, Hsiao C. A cytogenetic basis for transferring Russian wild rye from Elymus to Psathyrostachys[J]. Crop Science, 1983, 23(1): 123-126.[4] Baden C. A taxonomic revision of Psathyrostachys (Poaceae)[J]. Nordic Journal of Botany, 1991, 11(1): 3-26.[5] 郭本兆. 中国植物志(第九卷第三分册)[M]. 北京: 科学出版社, 1987.[6] Dewey D R. The genome system of classification as a guide to intergeneric hybridization with the perennial Triticeae[A].In: Gustafson J P. Gene Manipulation in Plant Improvement[M]. New York: Plenum Press, 1984: 209-279.[7] Hsiao C, Richard R, Wang C, et al. Karyotype analysis and genome relationships of 22 diploid species in the tribe Triticeae[J]. Canadian Journal of Genetics and Cytology, 1986, 28(1): 109-120.[8] Linde-Laursen I, von Bothmer R. Comparison of the karyotype of Psathyrostachys juncea and P. huashanica (Poaceae) studied by banding techniques[J]. Plant Systematics and Evolution, 1986, 151(3): 203-213.[9] Wang R C. Synthetic and natural hybrids of Psathyrostachys huashanica[J]. Genome, 1987, 29(6): 811-816.[10] 王秀娥, 李万隆, 刘大钧. 新麦草属两物种的C-分带研究[J]. 南京农业大学学报, 1998, 21(1): 10-13.[11] 孙根楼, 颜济, 杨俊良. 华山新麦草同高加索鹅观草和糙毛仲彬草间物种生物学研究[J]. 草业学报, 1995, 4(4): 39-45.[12] 刘芳, 孙根楼, 颜济, 等. 普通小麦和华山新麦草及其属间杂种F1同功酶分析[J]. 作物学报, 1992, (3): 169-175.[13] Zhang H Q, Zhou Y H. Meiotic pairing behaviour reveals differences in genomic constitution between Hystrix patula and other species of the genus Hystrix Moench (Poaceae, Triticeae)[J]. Plant Systematics and Evolution, 2006, 258(3): 129-136.[14] Zhang H Q, Yang R W, Dou Q W, et al. Genome constitutions of Hystrix patula, H. duthiei ssp.duthiei and H. duthiei ssp. longearistata (Poaceae: Triticeae)revealed by meiotic pairing behavior and genomic in-situ hybridization[J]. Chromosome Research, 2006, 14(6): 595-604.[15] 王丽, 杨娟, 郭晶, 等. 用RAPD检测华山新麦草自然居群的遗传结构和居群分化[J]. 生态学报, 2005, 25(4): 720-727.[16] 赵利锋, 李珊, 潘莹, 等. 华山新麦草自然居群沿海拔梯度的遗传分化[J]. 西北植物学报, 2001, 21(3): 391-400.[17] 刘占林, 李珊, 阎桂琴, 等. 华山新麦草自然居群的遗传结构和种内遗传多态性研究[J]. 遗传学报, 2001, 28(8): 769-777.[18] 刘占林, 宋颐, 李珊, 等. 华山新麦草开花物候期观测和自然种群基因流的间接估测[J]. 植物生态学报, 2001, 25(4): 426-430.[19] 岳明, 张林静, 马凯, 等. 华山新麦草濒危原因及种群繁殖对策[J]. 生态学报, 2001, 21(8): 1314-1320.[20] 王丽, 赵桂仿. 华山新麦草胚和胚乳的发育研究[J]. 西北植物学报, 2002, 22(6): 786-790.[21] 张林静, 王丽, 李智选, 等. 华山新麦草小孢子发生及雄配子体的形成[J]. 西北大学学报(自然科学版), 2002, 32(2): 77-80.[22] 曾亮, 袁庆华, 王方, 等. 冰草属植物种质资源遗传多样性的ISSR分析[J]. 草业学报, 2013, 22(1): 260-267.[23] 李杰勤, 王丽华, 詹秋文, 等. 20个黑麦草品系的SRAP遗传多样性分析[J]. 草业学报, 2013, 22(2): 158-164.[24] 黄晓辉, 胡小文, 徐宗海, 等. 羊草种子休眠和萌发的激素调控研究[J]. 草业学报, 2013, 22(5): 183-189.[25] 王美南, 商鸿生. 华山新麦草对小麦全蚀病菌的抗病性研究[J]. 西北农业大学学报, 2000, 28(6): 69-71.[26] 井金学, 傅杰, 袁红旭. 三个小麦野生近缘种抗条锈性传递的初步研究[J]. 植物病理学报, 1999, 2(2): 147-150.[27] 万永芳, 颜济, 杨俊良, 等. 小麦近缘野生植物的赤霉病抗性研究[J]. 植物病理学报, 1997, 27(2): 107-111.[28] 李庆, 叶华智. 小麦近缘野生植物对禾谷缢管蚜的抗性研究[J]. 中国农业科学, 2002, 35(6): 719-723.[29] Song S, Tao Y, Zhang H W, et al. Psathyrostachys huashanica, a potential resource for resistance to Barley yellow dwarf virus-GAV[J]. European Journal of Plant Pathology, 2013, 137: 217-221.[30] Petersen G, Seberg O, Baden C. A phylogenetic analysis of the genus Psathyrostachys (Poaceae) based on one nuclear gene, three plastid genes, and morphology[J]. Plant Systematics and Evolution, 2004, 249: 99-110.[31] 凡星, 张颖, 周永红. Psathyrostachys huashanica和Pseudoroegneria strigosa细胞色素c成熟蛋白亚基ccmFN基因的序列分析[J]. 四川农业大学学报, 2006, 24(3): 252-255.[32] 董玉琛.小麦远缘杂交育种[A]. 21世纪小麦遗传育种展望——小麦遗传育种国际学术讨论会文集[C]. 北京: 中国农业科技出版社, 2001: 12-16.[33] 陈漱阳, 张安静, 傅杰. 普通小麦与华山新麦草的杂交[J]. 遗传学报, 1991, 18(6): 508-512.[34] 陈漱阳, 侯文胜, 张安静, 等.普通小麦-华山新麦草异附加系的选育及细胞遗传学研究[J]. 遗传学报, 1996, 23(6): 447-452.[35] 孙根楼, 颜济, 杨俊良. 普通小麦和华山新麦草属间杂种的产生及细胞遗传学研究[J]. 遗传学报, 1992, 19(4): 322-326.[36] 侯文胜, 张安静, 杨群慧, 等. 普通小麦-华山新麦草异代换系的选育及细胞遗传学研究[J]. 西北植物学报, 1997, 17(3): 368-373.[37] Kishii M, Dou Q, Garg M, et al. Production of wheat-Psathyrostachys huashanica chromosome addition lines[J]. Genes & Genetic Systems, 2010, 85(4): 281-286.[38] Wang Y, Xie Q, Yu K F, et al. Development and characterization of wheat-Psathyrostachys huashanica partial amphiploids for resistance to stripe rust[J]. Biotechnology Letters, 2011, 33(6): 1233-1238.[39] Wang Y, Kang H Y, Zhang H Q, et al. Distribution of chromosome numbers in BC2 and BC1F1 progenies derived from Triticum aestivum×Psathyrostachys huashanica[J]. Caryologia, 2011, 64(1): 67-74.[40] Wang Y, Yu K F, Xie Q, et al. Cytogenetic, genomic in situ hybridization (GISH) and agronomic characterization of alien addition lines derived from wheat-Psathyrostachys huashanica[J]. African Journal of Biotechnology, 2011, 10(12): 2201-2211.[41] Kang H Y, Zhang H Q, Fan X, et al. Morphological and cytogenetic studies on the hybrid between bread wheat and Psathyrostachys huashanica Keng ex Kuo[J]. Euphytica, 2008, 162(3): 441-448.[42] Kang H Y, Zhang H Q, Wang Y, et al. Comparative analysis of the homoeologous pairing effects of phKL gene in common wheat×Psathyrostachys huashanica Keng ex Kuo[J]. Cereal Research Communications, 2008, 36(3): 429-440.[43] Kang H Y, Wang Y, Sun G L, et al. Production and characterization of an amphiploid between common wheat and Psathyrostachys huashanica Keng ex Kuo[J]. Plant Breeding, 2009, 128(1): 36-40.[44] Kang H Y, Chen Q, Wang Y, et al. Molecular cytogenetic characterization of the amphiploid between bread wheat and Psathyrostachys huashanica[J]. Genetic Resources and Crop Evolution, 2010, 57(1): 111-118.[45] Kang H Y, Zhong M Y, Xie Q, et al. Production and cytogenetics of trigeneric hybrid involving Triticum, Psathyrostachys and Secale[J]. Genetic Resources and Crop Evolution, 2012, 59(3): 445-453.[46] Kang H Y, Zeng J, Xie Q, et al. Molecular cytogenetic characterization and stripe rust response of a trigeneric hybrid involving Triticum, Psathyrostachys, and Thinopyrum[J]. Genome, 2012, 55(5): 383-390.[47] 赵继新, 陈新宏, 王小利, 等. 普通小麦-华山新麦草异代换系的分子细胞遗传学研究[J]. 西北植物学报, 2004, 24(12): 2277-2281.[48] 武军, 赵继新, 陈新宏, 等. 普通小麦-华山新麦草衍生后代的细胞学特点及GISH分析[J]. 麦类作物学报, 2007, 27(5): 772-775.[49] 武军, 赵继新, 陈新宏, 等. 普通小麦-华山新麦草异附加系的SSR分析[J]. 河北农业大学学报, 2007, 30(5): 9-13.[50] Du W L, Wang J, Wang L M, et al. A novel SCAR marker for detecting Psathyrostachys huashanica Keng chromatin introduced in wheat[J]. Genetics and Molecular Research, 2013, 12(4): 4797-4806.[51] 赵继新, 武军, 程雪妮, 等. 普通小麦-华山新麦草1Ns二体异附加系的农艺性状和品质[J]. 作物学报, 2010, 36(9): 1610-1614.[52] Kang H Y, Wang Y, Fedak G, et al. Introgression of chromosome 3Ns from Psathyrostachys huashanica into wheat specifying resistance to stripe rust[J]. PLoS ONE, 2011, 6(7): e21802. [53] Wang Y, Yu K, Xie Q, et al. The 3Ns chromosome of Psathyrostachys huashanica carries the gene(s) underlying wheat stripe rust resistance[J]. Cytogenetic and Genome Research, 2011, 134(2): 136-143.[54] Du W L, Wang J, Lu M, et al. Characterization of a wheat-Psathyrostachys huashanica Keng 4Ns disomic addition line for enhanced tiller numbers and stripe rust resistance[J]. Planta, 2013, 239(1): 97-105. [55] Du W L, Wang J, Lu M, et al. Molecular cytogenetic identification of a wheat-Psathyrostachys huashanica Keng 5Ns disomic addition line with stripe rust resistance[J]. Molecular Breeding, 2013, 31(4): 879-888.[56] Du W L, Wang J, Pang Y H, et al. Isolation and characterization of a Psathyrostachys huashanica Keng 6Ns chromosome addition in common wheat[J]. PLoS One, 2013, 8: e53921.[57] Du W L, Wang J, Wang L M, et al. Development and characterization of a Psathyrostachys huashanica Keng 7Ns chromosome addition line with leaf rust resistance[J]. PLoS One, 2013, 8: e70879.[58] 李振岐, 曾士迈.中国小麦锈病[M].北京: 中国农业出版社, 2002: 370-373.[59] 王殿波, 王美南, 井金学, 等. 小麦远缘杂交后代对小麦全蚀病抗病性研究[J]. 西北植物学报, 2003, 23(9): 1617-1620.[60] 傅杰, 王美南, 赵继新, 等. 抗全蚀病小麦-华山新麦草中间材料H8911 的细胞遗传学研究与利用[J]. 西北植物学报, 2003, 23(12): 2157-2162.[61] 傅杰, 赵继新, 陈漱阳, 等. 小麦-华山新麦草抗全蚀病新种质的分子细胞遗传学研究[J]. 西北植物学报, 2003, 23(11): 1905-1909.[62] 魏芳勤, 武军, 赵继新, 等. 普通小麦和华山新麦草衍生系H9021对全蚀病抗性的遗传分析[J]. 麦类作物学报, 2009, 29(1): 153-156.[63] 杭焱, 金燕, 卢宝荣. 濒危植物华山新麦草(Psathyrostachys huashanica)的遗传多样性及其保护[J]. 复旦学报(自然科学版), 2004, 43(2): 260-266.[64] 刘佩, 杨敏娜, 周新力, 等. 普通小麦-华山新麦草易位系H9020-1-6-8-3抗条锈病基因的遗传分析和SSR标记[J]. 植物病理学报, 2008, 38(1): 104-107.[65] 曹张军, 王献平, 王美南, 等. 小麦背景中来自华山新麦草的抗条锈病基因的遗传学分析和分子标记[J]. 遗传学报, 2005, 32(7): 738-743.[66] 丁春邦, 周永红. 小麦与华山新麦草远缘杂交的受精和胚胎发育[J]. 四川农业大学学报, 1997, 15(1): 18-20.[67] 廖进秋, 康厚扬, 杨瑞武, 等. 中国春ph2b突变体×华山新麦草F1自交和回交一代细胞遗传学研究[J]. 西北植物学报, 2007, 27(3): 442-448. |
[1] | 王晓丽,凡星,张春3,沙莉娜,张海琴,周永红. 用nrDNA ITS序列探讨小麦族含StH基因组物种的系统发育[J]. 草业学报, 2009, 18(6): 82-90. |
阅读次数 | ||||||
全文 |
|
|||||
摘要 |
|
|||||